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Crossover corrects for offsets in retinal circuitry. At the midpoint of the traces the voltages of the OFF bipolar and ON amacrine cells become more positive. At the ganglion cell, the bipolar cell current becomes more inward, but the inhibitory current from the ON amacrine cell becomes more outward. The contrast signals are in phase, but the offset currents cancel so that the OFF ganglion cell voltage is unaffected by the offsets in bipolar and amacrine cell voltage.
