Figure 2.

The spatial relationship between multiple Golgi cells Note that no restriction is imposed to axons, which spread and overlap into the granular layer, that apical dendrites ramify in the molecular layer without specific orientation, and that basal dendrites ramify in the granular layer over a surface smaller than that occupied by the axon. In this figure, the fundamental features of the Golgi cell are captured. (Fragment of a (neonatal) cat cerebellar convolution (vertical section); Table XVII, Opera Omnia, Golgi, 1903). ‘The drawing is specifically made to show shape, disposition, ramification laws, localisation and relationships of the large gangliar cells of the granular layer. Protoplasmatic prolongations branch dichotomously in a very different way compared to Pukinje cells. The most distal extensions of the branches often reach the molecular layer peripheral limit. Nervous prolongations, with their fine and repeated subdivisions, form a complicated interlacement with the result that it is impossible to follow the course of single prolongations. Such interlacement does not seem to have borders either toward the inside of the granular layer or toward the molecular layer. Thus, several of these interlacements obviously mix up to form a complicated plexus.’ Translated from Opera Omnia (Golgi, 1903).