Schematic diagram summarizing the types of steps myoV takes. a, difference between a normal step (leading head bound to the 13th actin subunit) and the shorter step myoV takes before a back step (leading head bound to the 11th actin subunit). When viewed down the actin axis, the two lever arms are in the same plane for a normal step, whereas for a step before a back step, there is an azimuthal angle of ∼28° between the two lever arms. b, schematic of a back step. The lever arms in green show the possible binding positions of the new trailing head. During a back step, the tail position changes by 29 ± 14 nm, whereas that of the heads is 49 ± 27.