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CowN is required for N2 fixation in the presence of CO but not for CO-dependent growth. R. rubrum strains UR2 (WT), UR616 (ΔcooA::aacC1Ω), UR2216 (rcoM::aacC1Ωlinker), and UR2546 (cowN::aacC1Ω) were cultivated photosynthetically (PS, plates A to C) on medium supplemented with NH4Cl in the presence of 0.15% CO (initial level) (A) and on medium requiring N2 fixation without (B) or with (C) 0.15% CO. Plate D was incubated in the dark under conditions suitable for CO-dependent growth (9).
