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Juxtaposition of Urm1 as a prokaryotic sulfur carrier and a eukaryotic UBL. The shaded areas indicate overlapping aspects of the pathways, illustrating that Urm1 seems to use features common to both ubiquitin and MoaD for its activation and conjugation. The Urm1–E2 intermediate indicated in brackets is still hypothetical but may account for the hydroxylamine sensitivity of Urm1–protein conjugates formed in response to oxidative stress.
