GM1 gangliosides, located on the surface of the targeted cells, are the known ligands that facilitate binding and internalization of CT. N-acetylgalactosamine (GalNAc), N-acetylneuraminic acid (Neu5Ac), glucose (Glc), and galactose (Gal) residues comprise the pentasaccharide moiety of the GM1 ganglioside facilitate interaction with the B subunit of cholera toxin (CTB).