Effects of CO2 on krill development. (a) An embryo reared under current surface pCO2 developed into the limb bud stage; (b) an embryo reared at 2000 µatm pCO2. Embryonic development was disrupted during gastrulation with a larger portion of the ectoderm appearing irregular. (c) Comparison of embryonic development under 380, 1000 and 2000 µatm pCO2 conditions determined 7–10 days of spawning during 2009 experiments. Dark blue, hatched; light blue, limb formation; yellow, blastula–gastrula; pink, cleavage; red, no development. Scale bar, (a,b) 0.6 mm.