TABLE 3.
Gene ID | Gene name | Annotation | ΔhrrA/wild-type mRNA ratiob |
|
---|---|---|---|---|
Heme | FeSO4 | |||
Two-component systems | ||||
cg3247 | hrrA | Two-component system, response regulator | 0.02 | 0.02 |
cg2200 | cgtR8 | Two-component system, response regulator | 1.79 | 2.13 |
cg2201 | cgtS8 | Two-component system, signal transduction histidine kinase | 1.99 | 2.42 |
Heme transport/utilization | ||||
cg0466 | htaA | Secreted heme transport-associated protein | 0.46 | 1.02* |
cg0467 | hmuT | Hemin-binding periplasmic protein precursor | 0.57 | 1.03* |
cg0468 | hmuU | Hemin transport system, permease protein | 0.60 | 1.04* |
cg0469 | hmuV | Hemin transport system, ATP-binding protein | 0.86* | 1.24* |
cg0470 | htaB | Secreted heme transport-associated protein | 9.00 | 8.71* |
cg0471 | htaC | Secreted heme transport-associated protein | 8.06 | 5.09 |
cg2202 | hrtB | ABC-type transport system, permease component | 10.90 | 40.79 |
cg2204 | hrtA | ABC-type transport system, ATPase component | 10.47 | 43.93 |
cg2445 | hmuO | Heme oxygenase | 0.06 | 0.12 |
cg3156 | htaD | Secreted heme transport-associated protein | 0.26 | 0.63* |
Heme-containing proteins | ||||
cg0645 | cytP | Cytochrome P450 | 0.20 | 0.32 |
cg2403 | qcrB | Cytochrome b, membrane protein | 0.35 | 0.47 |
cg2404 | qcrA | Rieske iron-sulfur protein | 0.36 | 0.47 |
cg2405 | qcrC | Cytochrome c1 | 0.36 | 0.48 |
cg2406 | ctaE | Cytochrome aa3 oxidase, subunit 3 | 0.34 | 0.46 |
cg2780 | ctaD | Cytochrome aa3 oxidase, subunit 1 | 0.51 | 0.62 |
cg3141 | hmp | Flavohemoprotein | 2.74 | 1.95 |
cg3369 | Rieske-type iron-sulfur protein | 0.57 | 0.75* | |
Heme biosynthesis | ||||
cg0497 | hemA | Glutamyl-tRNA reductase | 1.77* | 1.44* |
cg0498 | hemC | Porphobilinogen deaminase | 1.62* | 1.19* |
cg0516 | hemE | Uroporphyrinogen decarboxylase | 1.87 | 1.99 |
cg0517 | hemY | Protoporphyrinogen oxidase | 1.90* | 2.04 |
cg0518 | hemL | Glutamate-1-semialdehyde 2,1-aminomutase | 1.77 | 2.10 |
cg0519 | Putative phosphoglycerate mutase | 1.59* | 1.74 | |
cg0520 | Secreted thiol-disulfide isomerase or thioredoxin | 1.51* | 1.70 | |
cg0522 | ccsA | Cytochrome c biogenesis protein membrane protein | 1.34* | 1.53 |
cg0523 | Membrane protein required for cytochrome c biosynthesis | 1.37* | 1.75 | |
cg0524 | ccsB | Cytochrome c assembly membrane protein | 1.39* | 1.90 |
cg1734 | hemH | Ferrochelatase precursor | 2.41* | 2.20 |
This table includes all genes with a putative heme/iron-related function that are differentially expressed in the ΔhrrA mutant. A list of all genes with a ≥2-fold-altered mRNA level is provided in Table S2 in the supplemental material. Genes and operons shown to be directly regulated by HrrA have all been included in this table, irrespective of their relative expression.
Average ratio of genes showing a ≥2-fold-altered mRNA level (ΔhrrA/wild-type mRNA ratio) of C. glutamicum cells grown on CGXII minimal medium with 2.5 μM heme or 2.5 μM FeSO4 as the iron source. The mRNA ratios represent average values from three independent DNA microarray experiments (P ≤ 0.05). *, P > 0.05.