In normal cells, ceramide and its binding protein, I2PP2A, which is the inhibitor for PP2A, are mostly in a 1:1 ratio. Therefore, it is believed to be the binding and inactivation of ceramide by I2PP2A that liberates the active form of PP2A, which, in turn, acts upon c-Myc, leading to the dephosphorylation and degradation. In cancer cells, elevated levels of I2PP2A were observed, which inhibits most of the available PP2A and results in stable (active) oncogenic c-Myc. The stable form of c-Myc can mediate tumor growth and cancer progression by upregulating expression of several oncogenes.
I2PP2A: Protein phosphatase 2A inhibitor 2; PP2A: Protein phosphatase 2A.