Six measures of reproductive fitness (mean ± s.e.) for female greater anis in communal breeding groups with respect to group size (two-pair, white diamonds, or three-pair, black diamonds) and position in the laying order (first-laying, A; second-laying, B or third-laying, C). (a) Total number of eggs laid, (b) number of eggs lost to ejection, (c) number of eggs incubated in the final clutch, (d) number of eggs that survived to hatching, (e) number of nestlings that survived to fledging and (f) number of fledglings produced per egg laid. Asterisks indicate statistically significant (p < 0.05) differences between females within groups of the same size. Comparisons between groups of different sizes, and details of statistical tests, are given in the text.