Flower phenotypes of osmads13-3 osmads3-4. A, One osmads13-3 flower with the removal of the lemma and the palea showing normal lodicules and stamens but its pistil with three stigmas. B, Longitudinal section of one osmads13-3 flower showing abnormal development of the ovule. C, One osmads3-4 flower with the removal of the lemma and the palea. D, Longitudinal section of one osmads3-4 flower showing normal development of the carpel and the ovule. E, One osmads13-3 osmads3-4 flower with the removal of the lemma and the palea showing inner organs. F, Longitudinal section of one osmads13-3 osmads3-4 flower showing the formation of higher order carpels. G, SEM observation of one osmads13-3 osmads3-4 flower primordium with the removal of the lemma and the palea. H, SEM observation of one osmads13-3 osmads3-4 flower primordium displaying the loss of determinacy in the center. I, SEM observation of one osmads13-3 flower primordium at stage Sp6. J, SEM observation of one osmads3-4 flower primordium at stage Sp6. K, SEM observation of one osmads13-3 osmads3-4 flower primordium at stage Sp6. As in J, the primordium of one ectopic organ is emerged. L, SEM observation of a wild-type flower at early stage Sp8 showing the termination of the floral meristem activity. M, SEM observation of one osmads13-3 osmads3-4 flower at the early stage Sp8 showing the remaining activity of the floral meristem. N, Closeup of M. The primordium of the primary carpel, the secondary carpel, and the floral meristem are indicated. O, Expression pattern of OSH1 in a wild-type flower at stage Sp8. P, Expression pattern of OSH1 in one osmads13-3 osmads3-4 flower at stage Sp8. Q to S, Floral diagrams of osmads13-3 (Q), osmads3-4 (R), and osmads13-3 osmads3-4 (S). ca, Capel; ii, inner integument; l-a, lodicule-anther mosaic organs; ll, lodicule-like structure; lo, lodicules; mtp, marginal tissue of the palea; oi, outer integument; ov, ovule; pc, primary carpel; sc, secondary carpel; st, stamen; tc, tertiary carpel. Bars = 1 mm in A, C, E, and G; 100 μm in B, D, F, and I to L; and 500 μm in H.