Left, when the levels of dopamine are high relative to adenosine and acetylcholine, cAMP/PKA activity is low in indirect-pathway MSNs, biasing excitatory synapses onto those cells towards LTD. Meanwhile, cAMP/PKA activity is high in direct-pathway MSNs, biasing excitatory synapses onto those cells towards LTP. Indirect-pathway LTD and direct pathway LTP are proposed to cause an active, motivated behavioral state. Right, when the levels of dopamine are low relative to adenosine and acetylcholine, cAMP/PKA activity is high in indirect-pathway MSNs, biasing excitatory synapses onto those cells towards LTP. Meanwhile, cAMP/PKA activity is low in direct-pathway MSNs, biasing excitatory synapses onto those cells towards LTD. Indirect-pathway LTP and direct pathway LTD are proposed to induce an inactive, apathetic behavioral state.