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Morphological analysis of compound mutants for Gli2 and Gli3 alleles. A and B, Gli3 mutant male (A) and female GT (B) exhibit normal sexually dimorphic development at E18.5. C, The GT of the Gli2+/−;Gli3−/− embryos shows a groove-like structure similar to the phenotype of the Gli2 homozygotes. D, The Gli2−/−;Gli3+/− embryos display drastic GT defects with underdeveloped glans, which does not contain the urethra.
