Tree of highest likelihood from the 28Sbe+wg dataset. This tree represents the preferred phylogenetic hypothesis of Harpalinae. Thick branches show clades supported by greater than 95% posterior probability from Bayesian inference. Colored branches are members of the lebiomorph assemblage color coded by tribe. Clade (a) represents Chlaenini + Panagaeini + Oodini group, (b) Orthogoniini + Graphipterini + Pseudomorphini clade, (c) Platynini, (d) Zuphiitae clade, (e) Lebiini, (f) Harpalini, (g) Pentagonicini +Odacanthini clade, (h) Lachnophorini + Calophaena, (i) clade comprised of pterostichines and relatives.