Fig. 1.

Sequence and timing of trial events. A: target selection task. Trials began with subjects directing gaze at a central fixation point. The fixation point transiently turned into a luminance cue. After a randomized delay, the fixation point was extinguished and the stimuli appeared. If the stimuli were moving, the monkey had to smoothly pursue the target that matched the luminance cue; if they were stationary, the monkey had to make a saccade to the correct target. B: pursuit-only task. Trials began with subjects directing gaze to a central fixation point. After a randomized delay, a single target appeared and moved in a step-ramp fashion toward the fixation point. The monkey was rewarded for smoothly pursuing the target.