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GRAF1 up-regulation precedes expression of skeletal differentiation markers. C2C12 cells maintained in growth (G) media or exposed to LSM for the indicated times were fixed and stained with GRAF1 (green) and indicated skeletal muscle differentiation marker (red; Myg (myogenin); Tm (tropomyosin); MHC (myosin heavy chain)). Nuclei were labeled with DAPI (blue). Scale bar, 50 μm. Arrowheads highlight GRAF recruitment to bi-polar tips of differentiated myoblasts (also see Fig. 7A).
