(I) Strict consensus tree of 15 parsimonious trees inferred from petA-psbJ, psbD-trnT, trnT-trnE, and trnT-trnL spacer sequences. Numbers above branches refer to bootstrap support in parsimony analysis, and those below branches are Bayesian posterior probability. CI = 0·809, RI = 0·901. Roman numerals following species names represent the inversion type in the psbD-trnT region. (II) A penalized likelihood chronogram of the Bayesian consensus tree of the combined data. The star indicates the calibration point used (Lavin et al., 2005, for details, see Materials and methods), The time scale is in million years ago (Mya).