Table 1.
Strain | Species/complexa | Source/organ | Additional informationc | STb | Alleleb |
|||||
---|---|---|---|---|---|---|---|---|---|---|
gyrB | groL | gltA | metG | ppsA | recA | |||||
Reference/type strains | ||||||||||
CECT 4199T | A. allosaccharophila/A. veronii | Anguilla anguilla (eel) | 13 | 14 | 13 | 14 | 13 | 13 | 13 | |
NCIMB 1134 | A. bestiarum/A. hydrophila | Rainbow trout | 4 | 5 | 4 | 5 | 4 | 4 | 4 | |
DSM 13956T | A. bestiarum | Infected fish | 8 | 9 | 8 | 9 | 8 | 8 | 8 | |
CECT 838T | A. caviae/A. punctata subsp. caviae | Epizootic of young guinea pigs | 12 | 13 | 12 | 13 | 12 | 12 | 12 | |
NCIMB 882 | A. caviae | Goldfish (Crassius auratus) | 3 | 4 | 3 | 4 | 3 | 3 | 3 | |
DSM 11577T | A. encheleia | Healthy eel in freshwater | 7 | 8 | 7 | 8 | 7 | 7 | 7 | |
CECT 4487T | A. enteropelogenes | Human feces | 18 | 20 | 19 | 20 | 19 | 17 | 19 | |
CECT 4255T | A. enteropelogenes/A. trota | Human feces | 16 | 18 | 17 | 18 | 17 | 16 | 17 | |
DSM 17534T | A. eucrenophila | Freshwater fish | 9 | 10 | 9 | 10 | 9 | 9 | 9 | |
CECT 4228T | A. jandaei | Feces from patient with diarrhea | 14 | 15 | 14 | 15 | 14 | 14 | 14 | |
ATCC 7966T | A. hydrophila | Milk | 1 | 1 | 1 | 1 | 1 | 1 | 1 | |
CECT 398 | A. hydrophila | Human feces of a child with diarrhea | 11 | 12 | 11 | 12 | 11 | 11 | 11 | |
DSM 4881T | A. media | Fish farm effluent | 6 | 7 | 6 | 7 | 6 | 6 | 6 | |
DSM 19604T | A. popoffii | Drinking water | 10 | 11 | 10 | 11 | 10 | 10 | 10 | |
NCIMB 1109 | A. salmonicida subsp. achromogenes | Diseased sea trout, Salmo trutta | 3 | 2 | 3 | 2 | 0 | 2 | ||
NCIMB 2020 | A. salmonicida subsp. masoucida | Masou, Oncorhynchus sp. | 2 | 2 | 3 | 2 | 0 | 2 | ||
NCIMB 1102T | A. salmonicida subsp. salmonicida | Atlantic salmon | 2 | 2 | 2 | 2 | 2 | 2 | 2 | |
DSM 17445T | A. sharmana | Warm spring | 82 | |||||||
CECT 4240T | A. schubertii | Forehead abscess | 15 | 16 | 15 | 16 | 15 | 15 | 15 | |
CECT 4245T | A. sobria | Fish | 19 | 21 | 20 | 21 | 20 | 18 | 20 | |
NCIMB 75 | A. sobria | Diseased freshwater fish | 5 | 6 | 5 | 6 | 5 | 5 | 5 | |
CECT 4257T | A. veronii bv. veronii | Sputum of drowning victim | 17 | 19 | 18 | 19 | 18 | 16 | 18 | |
CECT 4246 | A. veronii bv. sobria | Infected frog (red leg disease) | 17 | 16 | 17 | 16 | 0 | 16 | ||
Field strains | ||||||||||
Ae1 | A. sobria-A. veronii | European catfish (Ameiurus melas)/kidney | H1W2S1 | 20 | 22 | 21 | 22 | 21 | 19 | 20 |
Ae2 | A. hydrophila | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S1 | 21 | 23 | 22 | 23 | 22 | 20 | 21 |
Ae3 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S2 | 22 | 24 | 23 | 24 | 23 | 21 | 22 |
Ae4 | A. sobria-A. veronii | Sea bass (Dicentrarchus labrax)/kidney | H3W2S2 | 23 | 25 | 24 | 25 | 24 | 22 | 23 |
Ae5 | A. sobria-A. veronii | Sturgeon (Acipenser sp.)/kidney | H1W2S2 | 24 | 26 | 25 | 26 | 25 | 23 | 24 |
Ae6 | A. hydrophila | Northern pike (Esox lucius)/kidney | H1W2S3 | 25 | 27 | 26 | 27 | 26 | 24 | 25 |
Ae7 | A. sobria-A. veronii | Trout (Salmo trutta)/kidney | H1W1S3 | 26 | 28 | 27 | 28 | 27 | 25 | 26 |
Ae8 | A. sobria-A. veronii | European catfish (Ameiurus melas)/kidney | H1W2S3 | 27 | 29 | 28 | 17 | 21 | 26 | 27 |
Ae9 | A. hydrophila | Crayfish (Procambarus clarkii)/hemolymph | H1W2S4 | 28 | 30 | 29 | 29 | 28 | 27 | 28 |
Ae10 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S3 | 29 | 31 | 30 | 30 | 29 | 28 | 29 |
Ae11 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S4 | 30 | 24 | 31 | 31 | 30 | 29 | 30 |
Ae12 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S4 | 31 | 25 | 32 | 32 | 31 | 3o | 31 |
Ae13 | A. sobria-A. veronii | European grayling (Thymallus thymallus)/kidney | H1W1S4 | 32 | 26 | 33 | 33 | 32 | 31 | 33 |
Ae14 | A. sobria-A. veronii | Arctic char (Salvelinus alpinus)/kidney | H1W1S4 | 33 | 32 | 34 | 34 | 33 | 32 | 34 |
Ae15 | A. sobria-A. veronii | Arctic char (Salvelinus alpinus)/kidney | H1W1S4 | 34 | 33 | 35 | 35 | 34 | 33 | 35 |
Ae16 | A. hydrophila | Rainbow trout (Oncorhynchus mikiss)/kidney | H1W1S4 | 35 | 34 | 36 | 36 | 35 | 34 | 36 |
Ae17 | A. sobria-A. veronii | Trout (Salmo trutta)/kidney | H1W1S1 | 36 | 35 | 37 | 37 | 36 | 35 | 37 |
Ae18 | A. sobria-A. veronii | Eel (Anguilla anguilla)/kidney | H2W2S4 | 37 | 36 | 38 | 38 | 37 | 36 | 38 |
Ae19 | A. hydrophila | Sea bream (Sparus aurata)/kidney | H3W2S1 | 2 | 2 | 2 | 2 | 2 | 2 | 2 |
Ae20 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S1 | 38 | 37 | 39 | 39 | 38 | 37 | 39 |
Ae21 | A. sobria-A. veronii | Goldfish (Carassius auratus)/spleen | H1W2S2 | 39 | 38 | 40 | 40 | 39 | 38 | 40 |
Ae22 | A. caviae-A. media | Sturgeon (Acipenser sp.)/kidney | H1W2S2 | 40 | 39 | 41 | 41 | 40 | 39 | 41 |
Ae23 | A. sobria-A. veronii | Goldfish (Carassius auratus)/kidney | H1W2S3 | 41 | 40 | 42 | 42 | 41 | 40 | 42 |
Ae24 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S3 | 42 | 41 | 43 | 43 | 42 | 41 | 43 |
Ae25 | A. sobria-A. veronii | Goldfish (Carassius auratus)/spleen | H1W2S3 | 43 | 42 | 44 | 44 | 43 | 42 | 44 |
Ae26 | A. sobria-A. veronii | Arctic char (Salvelinus alpinus)/liver | H1W1S3 | 44 | 43 | 45 | 45 | 27 | 43 | 45 |
Ae27 | A. hydrophila | Arctic char (Salvelinus alpinus)/spleen | H1W1S3 | 45 | 44 | 46 | 46 | 44 | 44 | 46 |
Ae28 | A. caviae-A. media | Goldfish (Carassius auratus)/spleen | H1W2S3 | 46 | 45 | 47 | 47 | 45 | 45 | 47 |
Ae29 | A. sobria-A. veronii | Sturgeon (Acipenser sp.)/kidney | H1W2S3 | 47 | 46 | 48 | 48 | 46 | 46 | 48 |
Ae30 | A. sobria-A. veronii | Channel catfish (Ictalurus punctatus)/kidney | H1W2S3 | 48 | 47 | 49 | 49 | 47 | 47 | 49 |
Ae31 | A. hydrophila | Trout (Salmo trutta)/kidney | H1W1S3 | 2 | 2 | 2 | 2 | 2 | 2 | 2 |
Ae32 | A. sobria-A. veronii | European catfish (Ameiurus melas)/liver | H1W2S3 | 49 | 48 | 50 | 50 | 48 | 48 | 50 |
Ae33 | A. sobria-A. veronii | European catfish (Ameiurus melas)/liver | H1W2S3 | 50 | 49 | 51 | 51 | 49 | 49 | 51 |
Ae34 | A. sobria-A. veronii | Carp (Cyprinus carpio)/liver | H1W2S3 | 51 | 50 | 52 | 50 | 50 | 50 | 52 |
Ae35 | A. sobria-A. veronii | European catfish (Ameiurus melas)/kidney | H1W2S4 | 52 | 51 | 53 | 52 | 51 | 51 | 53 |
Ae36 | A. sobria/A. veronii | Rainbow trout (Oncorhynchus mikiss)/kidney | H1W1S4 | 53 | 52 | 54 | 31 | 52 | 52 | 54 |
Ae37 | A. hydrophila | Carp (Cyprinus carpio)/kidney | H1W2S4 | 54 | 14 | 55 | 14 | 53 | 53 | 13 |
Ae38 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mikiss)/kidney | H1W1S4 | 55 | 53 | 56 | 53 | 54 | 54 | 55 |
Ae39 | A. sobria-A. veronii | Sea bream (Sparus aurata)/kidney | H3W2S1 | 56 | 54 | 57 | 54 | 55 | 55 | 56 |
Ae40 | A. sobria-A. veronii | Trout (Salmo trutta)/kidney | H1W1S1 | 38 | 37 | 39 | 39 | 38 | 37 | 39 |
Ae41 | A. hydrophila | Trout (Salmo trutta)/kidney | H1W1S1 | 57 | 9 | 58 | 55 | 8 | 56 | 57 |
Ae42 | A. sobria-A. veronii | Goldfish (Carassius auratus)/kidney | H1W2S1 | 58 | 55 | 59 | 56 | 56 | 57 | 58 |
Ae43 | A. hydrophila | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S1 | 59 | 56 | 60 | 57 | 57 | 58 | 59 |
Ae44 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S1 | 60 | 57 | 61 | 58 | 58 | 59 | 60 |
Ae45 | A. hydrophila | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S1 | 38 | 37 | 39 | 39 | 38 | 37 | 39 |
Ae46 | A. hydrophila | Trout (Salmo trutta)/kidney | H1W1S1 | 2 | 2 | 2 | 2 | 2 | 2 | 2 |
Ae47 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S1 | 61 | 58 | 62 | 59 | 59 | 60 | 61 |
Ae48 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S2 | 62 | 38 | 63 | 60 | 60 | 61 | 62 |
Ae49 | A. sobria-A. veronii | European perch (Perca fluviatilis)/kidney | H1W2S2 | 63 | 59 | 64 | 61 | 61 | 62 | 63 |
Ae50 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S2 | 64 | 60 | 65 | 62 | 62 | 63 | 64 |
Ae51 | A. hydrophila | Crayfish (Procambarus clarkii)/hemolymph | H1W2S3 | 65 | 61 | 66 | 63 | 63 | 64 | 65 |
Ae52 | A. sobria-A. veronii | Crayfish (Procambarus clarkii)/hemolymph | H1W2S3 | 66 | 62 | 67 | 64 | 64 | 65 | 66 |
Ae53 | A. sobria-A. veronii | Carp (Cyprinus carpio)/kidney | H1W2S3 | 67 | 38 | 40 | 40 | 39 | 66 | 40 |
Ae54 | A. sobria-A. veronii | Crayfish (Procambarus clarkii)/hemolymph | H1W2S3 | 68 | 63 | 68 | 65 | 65 | 67 | 67 |
Ae55 | A. sobria-A. veronii | Manila clam (Ruditapes philippinarum)/foot | H3W2S2 | 69 | 64 | 69 | 66 | 66 | 68 | 68 |
Ae56 | A. sobria-A. veronii | Freshwater mussel (Anodonta sp.)/foot | H1W2S3 | 70 | 65 | 70 | 67 | 67 | 69 | 69 |
Ae57 | A. sobria-A. veronii | Sea bass (Dicentrarchus labrax)/kidney | H3W2S1 | 71 | 66 | 71 | 68 | 68 | 70 | 70 |
Ae58 | A. hydrophila | Carp (Cyprinus carpio)/kidney | H1W2S1 | 72 | 67 | 72 | 69 | 69 | 71 | 71 |
Ae59 | A. sobria-A. veronii | Sea bass (Dicentrarchus labrax)/kidney | H3W2S3 | 23 | 25 | 24 | 25 | 24 | 22 | 24 |
Ae60 | A. sobria-A. veronii | Eel (Anguilla anguilla)/kidney | H2W2S3 | 73 | 68 | 73 | 70 | 70 | 72 | 11 |
Ae61 | A. hydrophila | Flathead gray mullet (Mugil cephalus)/kidney | H2W2S4 | 74 | 69 | 74 | 71 | 71 | 73 | 72 |
Ae62 | A. hydrophila | Sea bream (Sparus aurata)/kidney | H3W2S1 | 75 | 70 | 75 | 72 | 72 | 74 | 73 |
Ae63 | A. hydrophila | Rainbow trout (Oncorhynchus mykiss)/kidney | H1W1S2 | 76 | 71 | 76 | 73 | 73 | 75 | 74 |
Ae64 | A. sobria-A. veronii | Chub (Leuciscus cephalus)/skin | H2W2S2 | 77 | 66 | 77 | 74 | 74 | 76 | 75 |
Ae65 | A. hydrophila | Carp (Cyprinus carpio)/skin | H1W2S2 | 78 | 72 | 78 | 75 | 44 | 77 | 76 |
Ae66 | A. sobria-A. veronii | Carp (Cyprinus carpio)/skin | H1W2S2 | 79 | 73 | 79 | 76 | 75 | 78 | 77 |
Ae67 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mykiss)/brain | H1W1S2 | 80 | 14 | 80 | 14 | 13 | 79 | 78 |
Ae68 | Atypical | Rainbow trout (Oncorhynchus mykiss)/brain | H1W1S3 | 81 | 74 | 81 | 77 | 76 | 80 | 79 |
Ae69 | A. sobria-A. veronii | Trout (Salmo trutta)/kidney | H1W1S3 | 82 | 75 | 82 | 78 | 20 | 81 | 80 |
Ae70 | A. sobria-A. veronii | Trout (Salmo trutta)/kidney | H1W1S3 | 83 | 76 | 83 | 39 | 77 | 82 | 81 |
Ae71 | A. sobria-A. veronii | European perch (Perca fluviatilis)/kidney | H1W2S3 | 84 | 77 | 84 | 79 | 78 | 83 | 82 |
Ae72 | A. hydrophila | European catfish (Ameiurus melas)/liver | H1W2S4 | 85 | 78 | 85 | 80 | 79 | 84 | 83 |
Ae73 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mykiss)/skin | H1W1S4 | 86 | 79 | 86 | 81 | 80 | 85 | 84 |
Ae74 | A. sobria-A. veronii | Sea bream (Sparus aurata)/brain | H3W2S3 | 87 | 66 | 87 | 82 | 81 | 86 | 85 |
Ae75 | A. sobria-A. veronii | European perch (Perca fluviatilis)/kidney | H1W2S3 | 87 | 66 | 87 | 82 | 81 | 86 | 85 |
Ae76 | A. sobria-A. veronii | Rainbow trout (Oncorhynchus mykiss)/eyes | H1W1S3 | 88 | 80 | 88 | 83 | 82 | 87 | 86 |
Ae77 | A. sobria-A. veronii | Tench (Tinca tinca)/spleen | H1W2S4 | 89 | 81 | 89 | 84 | 83 | 88 | 87 |
The phenotypic group names (i.e., complexes, which are shown for field isolates) were assigned according to the proposed attribution provided by Khajanchi et al. (32), using the keys of identification previously described (1, 13).
STs and alleles were determined by MLST.
H, habitat (1, freshwater; 2, brackish water; 3, marine water); W, water (1, cold water; 2, warm water); S, season (1, winter; 2, spring; 3, summer; 4, autumn).