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Possible functions of TBS-I–III in the establishment of chromosome bi-orientation. The possible actions of TBS-I–III with Sgo1 or Htz1 in chromosome bi-orientation (bi-polar attachment) are described in Discussion. The nucleosome may be ‘the shugoshin (guardian) of shugoshin: Shugomaster’ to prevent its degradation by currently unknown proteases. The APC/C proteosomal complex is one candidate (Salic et al, 2004; Karamysheva et al, 2009).
