(A) The structure of N-ColE7 in complex with DNA. Zn2+ (in green sphere), two β-strands, and one α-helix compose the ββα-Me fold at the active site. (B) Definition of the positional coordinate of the protein (r, z, ϕ) in a cylindrical coordinate system and the orientational coordinate of the protein (rp, θp, ϕp) in a spherical coordinate system.