18Δm and 18Δp are measured and predicted 18O discrimination (‰) when θeq = 1 and θeq = 0.6 (full or partial isotopic equilibrium, respectively), ε = Ccs/(Ca − Ccs) where Ca and Ccs indicate ambient and chloroplast surface CO2 concentrations, Δea is isotopic discrimination between atmospheric CO2, δa, and that in equilibrium with leaf water, δe: Δea = 1,000 × [(δe/1,000 + 1)/( δa/1,000 + 1) − 1]. Irradiance values are in μmol photon m−2 s−1. Analysis and definitions are based on Gillon and Yakir (2001).