Table 2.

Lymnaeid shell measurement comparison between Lymnaea cousini, L. bogotensis (= L. cousini), L. ubaquensis (= Pseudosuccinea columella) and L. meridensis n. sp. (= L. cousini from Laguna Mucubaji, Venezuela), from their respective type localities in northern Andean countries

	L. cousini Chanchu-Yacu, Chillogallo Ecuador		L. bogotensis Savannah of Bogota Colombia		L. ubaquensis Laguna de Ubaque Colombia	L. meridensis n. sp. Mucubaji, Mérida State Venezuela
Shell parameters	Jousseaume (1887)1 n = n.s.	Paraense (1995)2 n = 24	Velasquez (2006)3 n = 30	present study n = 30	present study n = 30	present study n = 16
Shell length (SL)	10-14	6.6-8.5	3.1-11.7 (6.85 ± 2.3)	4.4-7.2 (6.03 ± 0.65)	7.9-11.8 (9.47 ± 0.97)	6.6-9.3 (8.05 ± 0.78)
Shell width (SW)	5-6/6-10	6.0	7.0	2.9-4.2 (3.64 ± 0.38)	4.6-6.9 (5.49 ± 0.52)	3.7-6.0 (5.24 ± 0.58)
Last spire length (LSL)	n.s.	n.s.	n.s.	3.9-6.3 (5.26 ± 0.56)	7.2-10.8 (8.62 ± 0.89)	6.1-8.4 (7.57 ± 0.13)
Spire length (SpL)	n.s.	3	4.6	1.3-2.5 (1.99 ± 0.26)	1.7-3.2 (2.37 ± 0.30)	1.6-2.7 (2.24 ± 0.31)
Aperture length (AL)	7-10	6	7.1	2.8-4.6 (3.79 ± 0.42)	5.5-8.6 (6.65 ± 0.72)	4.8-6.0 (5.70 ± 0.15)
Aperture width (AW)	4-6	4	5.3	1.9-3.0 (2.53 ± 0.28)	3.4-5.0 (4.06 ± 0.39)	2.5-3.9 (3.39 ± 0.13)
Whorl number	4	5	n.s.	3-4 (3.20 ± 0.41)	3-4 (3.13 ± 0.35)	3-3 (3.00 ± 0.00)
SW/SL ratio	n.s.	0.54-0.65 (0.59 ± 0.03)	0.54-0.71 (0.62 ± 0.05)	0.57-0.67 (0.60 ± 0.02)	0.53-0.65 (0.58 ± 0.03)	0.58-0.66 (0.62 ± 0.04)
AL/SL ratio	n.s.	0.61-0.69 (0.65 ± 0.02)	0.55-0.76 (0.64 ± 0.05)	0.60-0.67 (0.63 ± 0.02)	0.66-0.75 (0.70 ± 0.02)	0.65-0.73 (0.69 ± 0.04)
AL/SpL ratio	n.s.	1.59-2.23 (1.88 ± 0.18)	1.23-3.17 (1.84 ± 0.41)	1.62-2.19 (1.91 ± 0.15)	2.37-3.75 (2.82 ± 0.30)	2.17-3.03 (2.58 ± 0.43)
SpL/SL ratio	n.s.	0.31-0.38 (0.35 ± 0.02)	0.24-0.45 (0.36 ± 0.05)	0.30-0.37 (0.33 ± 0.02)	0.20-0.28 (0.25 ± 0.02)	0.24-0.30 (0.27 ± 0.03)

Range include minimum and maximum extremes, with mean and standard deviation SD in parentheses. Measurements in mm. n.s. = not specified. ¹ = [55]; ² = [40]; ³ = [43]