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. 2011 Sep 15;7(9):e1002283. doi: 10.1371/journal.pgen.1002283

Table 2. Relative mutation rates of nucleotide i to j per site i for ex-operonic intergenic sites were calculated from singleton SNPs for the two replicatory strands.

from A T C G Equilibrium frequency Equilibrium AT skew
Leading to A - 1.79965E-04 4.37222E-04 6.39046E-04 0.2257 -0.4176 (-0.6792, -0.1522)
T 2.34002E-04 - 1.67602E-03 1.17158E-03 0.5493
C 6.38189E-05 4.72409E-04 - 5.32538E-05 0.1319
G 6.59462E-04 4.49913E-05 0.00 - 0.0931
Lagging to A - 2.34003E-04 1.17158E-03 1.67602E-03 0.5493 0.4176 (0.6792, 0.1522)
T 1.79965E-04 - 6.39046E-04 4.37222E-04 0.2257
C 4.49913E-05 6.59462E-04 - 0.00 0.0931
G 4.72409E-04 6.38189E-05 5.32538E-05 - 0.1319

Relative rates were derived from the following leading strand ex-operonic SNP counts, where XY represents a change from nucleotide X to Y: AG 31 GA 12 CG 0 GC 1 GT 22 TA 8 TC 21 TG 2 AC 3 CA 6 AT 11 CT 23. Nucleotide frequencies at compositional equilibrium were derived from the relative mutation rates. Leading and lagging equilibrium AT skews were calculated from equilibrium A and T frequencies. 95% bootstrap intervals are shown in parentheses.