(A) Skeleton reconstruction of Edmontosaurus regalis. Scale bar, 100 cm. (B) Type skulls of the five named edmontosaur taxa: Thespesius edmontoni (CMN 8399), Edmontosaurus regalis (CMN 2288), E. saskatchewanensis (CMN 8509), E. annectens (YPM 2182, from [15]), Anatotitan copei (AMNH 5730). Scale bar, 20 cm. (C) Biostratigraphic distributions of edmontosaur species (in black) based on published synonymies. Grey bars indicate ghost ranges inferred by proposed taxonomic schemes. Abbreviations: Alta, Alberta; BS, Brett-Surman [35]; Camp, Campanian; Estd, Eastend Formation; Fr man, Frenchman Formation; HC, Hell Creek Formation; HWF, Horner et al. [13]; L Sco, Lower Scollard Formation; Lar, Laramie Formation; Lnc, Lance Formation; LW, Lull and Wright [15]; PM, Prieto-Márquez [36]; Sask, Saskatchewan; WH, Weishampel and Horner [14].