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Bees in the principal treatment groups (control, OA, and DA) all showed increase in learning index over time (Repeated measures MANOVA, time −10 blocks-: F = 13. 25; df = 9; P<0.001). OA group bees showed a significantly slower increase and reached a lower maximum learning index (Repeated measures MANOVA, groups (3): F = 4.06, df = 2, P<0.025).
