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. 2001 Apr 10;98(8):4534–4539. doi: 10.1073/pnas.071034898

Table 1.

Independent variables used in stepwise regression analyses of species richness

Independent variables Mnemonic Hypothesis
Precipitation (mm/yr−1) PREC A,B
Wet-day frequency (days) WET A,B
Mean annual temperature (C°) Tmean A,B
Mean daily maximum temperature (C°) Tmax A,B
Mean daily minimum temperature (C°) Tmin A,B
Mean daily temperature range (C°) Trange A,B
Mean vapor pressure (hPa) VAPOR A,B
Radiation (W/m2) RAD A,B
Cloud cover (Percent) CLOUD A,B
Frost frequency (days) FROST A,B
Mean wind speed (m/second) WIND A,B
Latitude LAT A,B,C
Quadrat area (km2) AREA A,B,C,D
Ecosystem diversity (number of ecosystems in quadrat) ECO A,C
Topographic relief (elevational range, m) TOPO A,D
Latitude × topographic relief L × T A,D

Letters under hypothesis refer to regression models to which variables have been assigned: (A) ad hoc model; (B) contemporary climate model (46, 13, 2830); (C) ecosystem diversity model (3, 5); and (D) topography × latitude model (1416, 31). All variables are correlated significantly with avian species richness (P < 0.001, Bonferroni-adjusted for simultaneously testing) at the 1° × 1° latitude-longitude resolution (n = 1,679 quadrats).