Table 2. Pairwise relatedness (mean r ± SE) of mature subordinate males, subordinates females and all subordinates combined with dominant females, dominant males and midpoint dominant pair (average r between the dominant pair).

	Midpoint r	Dominant males	Dominant females
Male subordinates (n = 13)	0.172±0.033	0.096±0.084	0.199±0.055
Female subordinates (n = 16)	0.070±0.067	0.018±0.084	0.122±0.081
Subordinates combined (n = 39)*	0.096±0.242	0.058±0.311	0.116±0.296

*Includes n = 10 subordinates with unclear sex.

All subordinates used to calculate relatedness were larger than 30 mm SL (mean SL: 38.7±5.4 mm; range: 30.0–48.5 mm SL).