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Relative reduction in hydrogenase activity of proton pathway variants. kobsrel (rate of the variant enzyme relative to that of the native enzyme) values for H2 evolution and uptake activities are shown. To determine the effect of substitution, the specific activity of the native enzyme was normalized to 1.0. Variant enzyme kobsrel values were calculated from individual experiments and averaged. Error bars denote S.D. (n = 3–8). AU, arbitrary units.
