1 | Gonopod tip like a single, very small, rounded lobule (Figs 5 & 6). Virtually no modifications between ♂ coxae 4 (Fig. 4H & I). Pantropical | Orthomorpha coarctata |
– | Gonopod tip mainly bi- to trifid. Modifications between ♂ coxae 4 mostly present. Local in Southeast Asia | 2 |
2 | Gonopod femorite about as long as prefemoral portion (Fig. 113) | Orthomorpha butteli |
– | Gonopod femorite ca 2–3 times longer than prefemoral portion (e.g. Figs 83, 94 & 99) | 3 |
3 | No distinct lateral sulcus on gonopod femorite demarcating a “postfemoral” region, at most a very faint one; solenophore tip neither acuminate nor branching. Thailand and northernmost Malaysia | 4 |
– | A distinct lateral sulcus on gonopod femorite demarcating a “postfemoral” region; solenophore tip either acuminate or rather evidently branching (e.g. Figs 30, 33 & 46) | 9 |
4 | At least surface of metaterga leathery, dull | 5 |
– | Dorsal surface shining | 7 |
5 | Only caudal parts of paraterga contrasting light. Paraterga less strongly developed, narrower, with only a small incision at best in front 1/3 extent. No tergal setae borne on evident knobs/tubercles | Orthomorpha sericata |
– | Entire paraterga contrastingly light, much broader, largely with an evident, dentiform incision in front 1/3 extent. At least some tergal setae borne on evident knobs/tubercles | 6 |
6 | Surface of metaterga rough, microgranulate and rugulose. Metatergal setigerous tubercles higher, present also on collum. Paratergal incisions especially prominent | Orthomorpha enghoffi |
– | Surface of metaterga nearly smooth, microgranulate. Metatergal setigerous tubercles evidently smaller, absent from collum. Paratergal incisions evident, but not so prominent | Orthomorpha alutaria |
7 | Adults > 30 mm long and > 4 mm wide. Collum broadly rounded laterally. Paraterga set high, at least some upturned above dorsum. Sternal cones between ♂coxae 4 isolated. Lateral sulcus on gonofemorite traceable, but very faint | Orthomorpha parasericata |
– | Adults < 30 mm long and < 4 mm wide. Collum narrowly rounded to angulate laterally. Paraterga set high, but remaining below dorsum even in male. Sternal cones between ♂ coxae 4 fused basally into a single lamina. Lateral sulcus on gonofemorite absent | 8 |
8 | Bases of paraterga more broadly light. Legs shorter, midbody ones ca 1.2 (♂) or 1.0 times (♀) as long as body height. Tip of solenophore like an elongate and undulate lobe | Orthomorpha subsericata |
– | Bases of paraterga more narrowly light. Legs longer, midbody ones ca 1.5 (♂) or 1.2–1.3 times (♀) as long as body height. Tip of solenophore tridentate | Orthomorpha asticta |
9 | Gonopod tip bifid (Figs 10, 12, 18 & 36) | 10 |
– | Gonopod tip trifid, albeit middle prong can be very small (Figs 44, 46, 52 & 61) | 18 |
10 | Metaterga smooth and shining, at most faintly rugulose (Figs 11 & 25) | 11 |
– | Metaterga at least with one rather evident, transverse row of tubercles near caudal margin (Figs 9, 13, 28 & 34) | 13 |
11 | Gonopod femorite slightly, but evidently twisted near base; tip rather deeply split, both terminal and subterminal prongs spiniform | Orthomorpha paviei |
– | Gonopod femorite not twisted near base; tip shallowly bifid, both terminal and subterminal prongs dentiform (e.g. Figs 12, 26 & 27) | 12 |
12 | Length ca 19–23 mm, width of midbody pro- and metazona 1.8 and 2.7 mm (♂), 2.3–2.5 and 3.4–3.5 mm (♀), respectively. Coloration of paraterga and epiproct creamy orange (live material) to faded pinkish or pale yellow (alcohol-preserved material) (Fig. 11). Similan Islands, Thailand (Map 2) | Orthomorpha picturata sp. n. |
– | Length 31–38 mm, width of midbody pro- and metazona 2.6–4.0 and 4.2–4.7 mm (♂), 3.2–3.8 and 4.8–5.3 mm (♀), respectively. Coloration of paraterga and epiproct creamy yellow (both live and alcohol-preserved material) (Fig. 25). Continental Thailand (Map 2) | Orthomorpha communis sp. n. |
13 | Metaterga with only a single row of very small tubercle near caudal margin (Figs 28, 31 & 34) | 14 |
– | Metaterga with two rows of tubercles: small ones in front of, larger ones behind sulcus (near caudal margin) (Figs 9, 13 & 22) | 16 |
14 | Caudal corners of anterior and midbody paraterga not extended behind tergal margin (Fig. 34A-D). Metaterga mostly flavous (Fig. 34A-F). Sternal cones between ♂ coxae 4 widely separated (Fig. 34H & I) | Orthomorpha suberecta sp. n. |
– | Caudal corners of all postcollum paraterga evidently produced behind tergal margin. Metaterga mostly infuscate. Sternal cones between ♂ coxae 4 subcontiguous | 15 |
15 | Pleurosternal carinae complete high crests with a sharp caudal tooth on segments 2–7 (♂). Transverse sulcus visible on metaterga 5–18, narrow, not reaching bases of paraterga, ribbed at bottom. Stricture between pro- and metazona narrow and rather shallow, faintly beaded at bottom down to base of paraterga (Fig. 34B-F & K) | Orthomorpha latiterga sp. n. |
– | Pleurosternal carinae complete high crests with a sharp caudal tooth on segments 2–4(5) (♂, ♀). Transverse sulcus evident (Fig. 28B-F & H), thin, deep and only slightly incomplete on metaterga 2–4, complete, at most very faintly beaded at bottom, reaching bases of paraterga on metaterga 5–18 (Fig. 28B, D & F) | Orthomorpha atypica sp. n. |
16 | A single sternal cone between ♂ coxae 4 (Fig. 9H & I). Vietnam | Orthomorpha arboricola |
– | Two sternal cones between ♂ coxae 4 (Figs 13H, I, 22H & I). Thailand | 17 |
17 | Postcollum paraterga with a very strong front indentation laterally (Fig. 22a-F & J-L) | Orthomorpha subtuberculifera sp. n. |
– | Postcollum paraterga with only a very small front indentation laterally (Fig. 13B-F) | Orthomorpha tuberculifera sp. n. |
18 | Paraterga mostly level to or even above dorsum | 19 |
– | All paraterga below dorsum | 28 |
19 | Colour pattern not strikingly contrasting, calluses only being inconspicuously paler than a dark remaining background (Figs 41 & 109) | 20 |
– | Coloration of metaterga more or less strongly contrasting, with very pale calluses against a very dark remaining background (Figs 49, 51 & 106) | 21 |
20 | Metaterga with two transverse rows of evident tubercles (Fig. 41A-F). Vietnam | Orthomorpha scabra |
– | Metaterga quite smooth, without transverse rows of evident tubercles (Fig. 109B-F). Northern Malaysia | Orthomorpha spiniformis sp. n. |
21 | Calluses on metaterga rather thin (Figs 51A-F & 100A-F) | 22 |
– | Calluses on metaterga rather thick (Fig. 49A-F) | 23 |
22 | Sternal cone between ♂ coxae 4 single, large (Fig. 51H & I). Southern Vietnam | Orthomorpha glandulosa |
– | Two small sternal cones between ♂ coxae 4 (Fig. 100I & J). Northern Malaysia | Orthomorpha elevata sp. n. |
23 | Metaterga virtually smooth, at most faintly rugulose, only seldom (Orthomorpha weberi) with minute knobs caudolaterally (Fig. 49A-F & J-L) | 24 |
– | Metaterga with a caudal row of evident tubercles | 26 |
24 | ♂ tarsal brushes entirely wanting. Epiproct with very distinct apical papillae. Lombok Island, Indonesia | Orthomorpha francisca |
– | ♂ tarsal brushes present. Epiproct with small apical papillae. Java Island, Indonesia | 25 |
25 | Tarsal brushes present until ♂ midbody legs | Orthomorpha weberi |
– | Tarsal brushes present until ♂ legs 7 | Orthomorpha tenuipes |
26 | Adult body particularly small, only ca 2.6 mm in width (♂). Bali Island, Indonesia | Orthomorpha baliorum |
– | Adult body larger, at least 3.5 mm in width. Java Island, Indonesia | 27 |
27 | Paraterga relatively strongly produced caudally, paratergal calluses in dorsal view rather strongly sinuate near ozopores | Orthomorpha conspicua |
– | Paraterga relatively slightly produced caudally, paratergal calluses in dorsal view rather poorly sinuate near ozopores | Orthomorpha beroni |
28 | A row of very small tubercles near caudal margin of some metaterga (Figs 39A, C, F, 43A, C, F, J-L) | 29 |
– | Metaterga virtually smooth, at best only slightly rugulose near caudal margin (Figs 53A-F, 58B-F) | 31 |
29 | Small tubercles present only on metaterga 10–18(19) | Orthomorpha flaviventer |
– | Small tubercles present on all postcollum metaterga (Figs 39A-F, 43A-F & J-L) | 30 |
30 | Paraterga more strongly produced caudally (Fig. 39A-G). Java Island, Indonesia | Orthomorpha unicolor |
– | Paraterga less strongly produced caudally (Fig. 43A-G & J-L). Vietnam | Orthomorpha rotundicollis |
31 | A single sternal lobe between ♂ coxae 41 (Fig. 53H & I) | 32 |
– | Two sternal cones, sometimes very low ones, between ♂ coxae 4 (Figs 65I, J, 81H, I, 106I & J) | 33 |
32 | Sternal lobe between ♂ coxae 4 subquadrate. Perak State, southern Malaysia | Orthomorpha bipunctata |
– | Sternal lobe between ♂ coxae 4 divided into two lobules only apically (Fig. 53H & I). Java Island, Indonesia | Orthomorpha zehntneri |
33 | Metaterga and collum usually more or less flavous, with lighter bands, spots or markings, at least in caudal parts (Figs 64A-F, 70A-F, 75A-G & K-M) | 34 |
– | Metaterga and collum between paraterga usually completely dark (Figs 37A-F, J-L, 106A-F) | 41 |
34 | Pore-bearing midbody paraterga particularly thick in lateral view (Figs 75E & 91E) | 35 |
– | Pore-bearing midbody paraterga considerably thinner in lateral view (Figs 65E, 69D & 86D) | 36 |
35 | Paraterga set mostly slightly higher (Figs 91E, H, 92E & H). Transverse sulcus present on metaterga 4–19. Tarsal brushes present until ♂ legs of segment 16 | Orthomorpha insularis |
– | Paraterga set mostly slightly lower (Figs 74B, D, E, 75C, E & H). Transverse sulcus present on metaterga 5–18(19). Tarsal brushes present until ♂ legs of segment 13 | Orthomorpha subkarschi |
36 | Entire collum yellow, colour pattern like in Figs 70, 87 or 97, with at least a small patch remaining light near caudal margin | Orthomorpha karschi |
– | Collum and subsequent tergites between paraterga at least partly infuscate, only postsulcus halves of metaterga often pallid to pale brown | 37 |
37 | No dark axial stripe on metaterga. Paraterga less prominent, their anterolateral margin relatively narrowly rounded (Figs 87 & 97) | 38 |
– | A dark axial stripe or line on metaterga evident. Paraterga much more prominent, their anterolateral margin very broadly rounded (Figs 65, 70 & 81) | 39 |
38 | Colour pattern as in Fig. 97. Calluses on paraterga narrow. Sternal cones between ♂ coxae 4 very small and broadly separated (Fig. 97H & I). Central Thailand | Orthomorpha isarankurai sp. n. |
– | Colour pattern as in Figs 86 & 87. Calluses on paraterga considerably broader. Sternal cones between ♂ coxae 4 much higher and poorly separated (Fig. 87G & H). Southern Thailand | Orthomorpha lauta |
39 | Sternal cones between ♂ coxae 4 smaller (Figs 81H, I, 82I & J). Caudal denticle of pleurosternal carinae traceable until ♂ segment 16 or 17 | Orthomorpha thalebanica |
– | Sternal cones between ♂ coxae 4 larger (Figs 64I & 70I). Caudal denticle of pleurosternal carinae traceable until ♂ segment 13 or 14 | 40 |
40 | Colour pattern of live animals as in Fig. 70A. Bases of paraterga in alcohol material mostly infuscate, brown to dark brown (Figs 69 & 70) | Orthomorpha horologiformis |
– | Colour pattern of live animals as in Fig. 65A. Bases of paraterga in alcohol material flavous, brown to dark brown (Figs 64 & 65) | Orthomorpha pterygota |
41 | Postcollum metaterga flavous behind transverse sulcus (Figs 79 & 103) | 42 |
– | Postcollum metaterga dark on both pre-sulcus and postsulcus halves (Figs 37, 58 & 106) | 45 |
42 | Paraterga with anterolateral corner considerably more angulate, more narrowly rounded | Orthomorpha fuscocollaris |
– | Paraterga with anterolateral corner considerably more broadly rounded (Figs 79 & 103) | 43 |
43 | Metaterga nearly completely flavous, only a thin axial line and transverse sulcus forming an infuscate cross. King Island, Mergui Archipelago, Myanmar | Orthomorpha crucifer |
– | Metaterga mostly dark, including bases of paraterga (Figs 79 & 103). Southern mainland Thailand | 44 |
44 | Pleurosternal carinae present at least as a caudal denticle until segment 12 (♂). Tarsal brushes present until about ♂ midbody legs | Orthomorpha banglangensis |
– | Pleurosternal carinae present at least as a caudal denticle until segment 16 (♂) or 13 (♀). Tarsal brushes present until about ♂ legs 8 | Orthomorpha subelevata sp. n. |
45 | Caudal corner of midbody calluses only modestly produced behind tergal margin (Figs 37C, D, K, 106D & E). Sternal cones between ♂ coxae 4 very small, nearly indistinguishable, widely separated (Figs 37H, I, 106I & J) | 46 |
– | Caudal corner of midbody calluses very strongly produced behind tergal margin (Figs 2A-F, 47C, D, 55C, D, 57C & D). Sternal cones between ♂ coxae 4 mostly evident, higher and less strongly separated (Figs 2H, I, 57H, I, 62H & I) | 47 |
46 | Pleurosternal carinae complete crests on segments 2–4 (♂, ♀) (Fig. 37B, D & E), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 17 (♂) or 16 (♀). Tarsal brushes present on ♂ legs 1–7 | Orthomorpha hydrobiologica |
– | Pleurosternal carinae complete crests bulged anteriorly and with a sharp caudal tooth on segments 2–7, thereafter only a sharp caudal tooth on segments 8–16 (♂), or crests bulged anteriorly and with a sharp caudal tooth on segments 2–4, thereafter only a small sharp caudal tooth on segments 5–14 (♀). Tarsal brushes present until legs of ♂ segment 13 | Orthomorpha similanensis sp. n. |
47 | Caudal corner of paraterga evidently rounded (Fig. 55A-F). Sternal cones between ♂ coxae 4 high and strongly separated (Fig. 55H & I). Environs of Kuala Lumpur, Malaysia | Orthomorpha fluminoris |
– | Caudal corner of paraterga evidently pointed (Figs 2A, C, F, 47A-G, J-L & 57A-G). Sternal cones between ♂ coxae 4 usually much lower, if not, then both fused at base (Figs 47H, I, 57H, I, 62H & I) | 48 |
48 | Paraterga especially strongly produced behind tergal margin (Figs 57 & 58). Sternal cones fused at base (Figs 57H, I, 58I & J). Singapore | Orthomorpha murphyi |
– | Paraterga much less strongly produced behind tergal margin (Figs 2A-G, 47A-G & 62A-G). Sternal cones not fused at base (Figs 2H, I, 47H, I, 62H & I) | 49 |
49 | Transverse metatergal sulcus present only on segments 5–18 (Fig. 2A, C & F). Pulau Karimunjawa and ?Java islands, Indonesia | Orthomorpha beaumontii |
– | An incomplete transverse metatergal sulcus traceable also either on segments 3 and 4 or only on segment 4 | 50 |
50 | Transverse metatergal sulcus traceable on segments 3 and 4, fully developed on segments 5–18 (Fig. 62A, C & F). Sternal cones between ♂ coxae 4 rather poorly separated (Fig. 62H & I). Sumatra Island, Indonesia | Orthomorpha melischi |
– | Transverse metatergal sulcus incomplete only on segment 4, complete on segments 5–18 (Fig. 47A, C, F & J-L). Sternal cones between ♂ coxae 4 strongly separated (Fig. 47H & I). Cambodia | Orthomorpha cambodjana |
1 Unknown in Orthomorpha fuscocollaris, a species based only on ♀♀.