Standard recruitment assays with proteins and T3 added as indicated were carried out as described in Experimental Procedures and bound proteins were detected by immunoblot.
(A). MED1-independent, T3-dependent binding of PGC-1α to the TRα-RXRα-UCP-1 enhancer complex.
(B). PGC-1α independent binding of MED1, as well as cooperative binding of intact MED1 and PGC-1α, to the TRα-RXRα-UCP-1 enhancer complex.
(C) and (D). Competitive binding of MED1(580–701) and PGC-1α to the TRα-RXRα-UCP-1 enhancer complex. All factors were added concommitantly to the immobilized DNA template. as indicated. In panel D the amount of PGC-1α in lane 4 was 10 times the amount in lane 3 and in the analyses in panel C
(E). Active displacement of prebound PGC-1α from the TRα-RXRα-UCP-1 enhancer complex by MED1(580–701). In lane 2 and 5, “1” and “2” indicate the order of addition of the factors to the preformed TRα-RXRα-UCP-1 enhancer complex. Factor 1 was added first and incubated for 30 minutes prior to addition of factor 2 and incubation for another 30 minutes