HeLa cells were transfected with MMP1, PRUNE, or UGT2B15 luciferase reporter vectors and an expression vector for ERα, with or without an expression vector for A-Fos. The cells were then treated with Ral (100 nM) or E2 (100 nM), as indicated, before determination of luciferase activity. Each bar represents the mean ± SEM for n ≥ 3 separate experiments. Bars marked with an asterisk are significantly different from the corresponding controls (Student’s t-test, p < 0.05).