Table 1.

Model parameters, their meanings and values used in the numerical simulations. For a sensitivity analysis of the model to parameters involved in resource dynamics, see electronic supplementary material, appendix S2.

parameter	meaning	value and range explored
TR+ TL	length of epoch	40
TR	length of productive season	1 to 40
S	basal resource supply	0.01
A	amount of resource during TR	0.1 to 10 (log scale)
ρR	resource degradation rate	0.1 (main text), 0.5
ρV	victim mortality rate	0.1 (main text), 0.2, 0.5
ρE	enemy mortality rate	0.1 (main text), 0.2, 0.5
ɛ	victim conversion rate	10
Φ0	basal attack rate	0.01
γ0	victim consumption rate	0.01
β0	progeny size (enemy)	3
∂	baseline cost of adaptation	0.4
w	non-optimal attack score	0.2
kV	victim cost of defence	0.25 (main text), 0.5
kE	enemy cost of attack	0.25 (main text), 0.5