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. 2011 Nov 29;2:86. doi: 10.3389/fphys.2011.00086

Figure 3.

Figure 3

Portrait of Herophilos (ca. 335–280 BC). He was the first to measure the heart beat using a water clock to time the pulse. Source: Reproduced with permission from the John P. McGovern Historical Collections and Research Center; Houston Academy of Medicine-Texas Medical Center Library; Houston, TX, USA. P-254, color photo; Artist: Joseph F. Doeve, painted in 1953.