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. 2011 Dec 2;6(12):e28554. doi: 10.1371/journal.pone.0028554

Table 1. Nineteen bioclimatic variables derived from the WorldClim database.

Abbreviation Variable Description
BIO1 Annual Mean Temp (C)
BIO2 Mean Diurnal Range (C)
BIO3 Isothermality (100 * BIO2 / BIO7)
BIO4 Temp Seasonality (100 * SD)
BIO5 Max Temp of Warmest Month (C)
BIO6 Min Temp of Coldest Month (C)
BIO7 Temp Annual Range (C) (BIO5–BIO6)
BIO8 Mean Temp of Wettest Quarter (C)
BIO9 Mean Temp of Driest Quarter (C)
BIO10 Mean Temp of Warmest Quarter (C)
BIO11 Mean Temp of Coldest Quarter (C)
BIO12 Annual Precip (mm)
BIO13 Precip of Wettest Month (mm)
BIO14 Precip of Driest Month (mm)
BIO15 Precip Seasonality (CV)
BIO16 Precip of Wettest Quarter (mm)
BIO17 Precip of Driest Quarter (mm)
BIO18 Precip of Warmest Quarter (mm)
BIO19 Precip of Coldest Quarter (mm)