Table 2.

Symbols used in the simulations and their descriptions.

Symbol	Description
F1	Foraging allele: pure producer (1.0 probability to apply producer strategy)
F2	Foraging allele: part-time scrounger (0.5 probability to apply producer strategy)
L0	Learning allele: non-learning
L1	Learning allele: complex learning rule
L2	Learning allele: simple learning rule
M1	Modifier of recombination allele: increasing recombination rate
M2	Modifier of recombination allele: decreasing recombination rate
S1	Learning switch allele: assigning learning rule at random
S2	Learning switch allele: assigning part-time scroungers with the complex learning mechanism, and pure producers with the simple learning mechanism
S3	Learning switch allele: assigning part-time scroungers with simple learning, and pure producers with complex learning
S4	Learning switch allele: assigning complex learning regardless of foraging genotype
S5	Learning switch allele: assigning simple learning regardless of foraging genotype
S*1	Foraging switch allele: assigning foraging strategy at random
S*2	assigning simple learners with the pure producer strategy, and complex learners with the part-time scrounger strategy
S*3	assigning simple learners with the part-time scrounger strategy, and complex learners with the pure producer strategy
S*4	assigning the pure producer strategy regardless of learning rule
S*5	assigning the part-time scrounger strategy regardless of learning rule
r	Recombination rate
r1	Recombination rate for parental genotype M1M2
r2	Recombination rate for parental genotype M1M2
r3	Recombination rate for parental genotype M2M2
R	Recombination rate between modifier locus and the foraging gene
n	Number of agents in the population
J	Number of steps in the game (equivalent to agents' lifespan)
G	Number of generations
μ	Mutation rate
γ	Cost of complex learning rule (% deducted from final cumulative payoff)
α	memory factor (weight of past events in memory when updating a new experience)