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Feedback and feedfoward circuits are fundamental building blocks of cortical inhibition. A, Feedback inhibition arises when cortical principal cells (grey) make excitatory synaptic contacts (red) on local interneurons (blue) that in turn form inhibitory synaptic contacts (blue triangles) on the principal cell population. B, Feedforward inhibition is generated when long-range excitatory afferent inputs (red) diverge onto both principal cells and local interneurons.
