The immutans variegation mutant of Arabidopsis.The green sectors contain normal-appearing chloroplasts, whereas the white sectors are heteroplastidic and contain abnormal plastids that lack pigments and lamellar structures, as well as rare normal chloroplasts (Wetzel et al., 1994). The white sectors accumulate the carotenoid precursor phytoene (Wetzel et al., 1994), and white sector formation is promoted by increased light intensity and temperature (Rédei, 1963; Röbbelen, 1968).