Table 1. Model comparison.

Model	Interaction zone		/rads					KL/bits	/bits	Consistent?
Null	None	−7.5	N/A	N/A	N/A	N/A	N/A	10.2	−944	No
MF	Global	−7.5	N/A	N/A	0.51	N/A	N/A	0.6	−863	Yes
T	K nearest-neighbours	−7.5	N/A	3.4	0.72	N/A	N/A	2.0	−864	No
S1	Spatial, symmetric	−7.5		N/A	1.42	N/A	N/A	1.3	−847	No
S2	Spatial, forward	−7.5		N/A	1.71	N/A	N/A	7.2	−834	No
S3	Spatial, symmetric	−7.5		N/A	1.80	−0.01	N/A	2.0	−856	No
S4	Spatial, forward	−7.5		N/A	1.74	0.20	N/A	2.3	−859	No
D1	Spatial, symmetric	−7.5		N/A	1.62	N/A	0.47	0.7	−827	Yes
D2	Spatial, forward	−7.5		N/A	1.50	N/A	0.46	1.5	−830	No
D3	Spatial, symmetric	−7.5		N/A	1.44	0.23	0.87	0.4	−828	Yes
D4	Spatial, forward	−7.5		N/A	1.44	0.37	0.82	0.8	−830	Yes

The interaction zone structure of each model, along with the (mean a posteri ) inferred values of model parameters, an indicator for the model's consistency with the observed large scale behaviour of the real system, The KL divergence between experimental and model results indicating large-scale goodness-of-fit and the log marginal likelihood () of the model calculated from the fine scale dynamics (as shown in Figure 3). N/A indicates that the model does not include the indicated parameter. The interaction zone indicates whether prawns interact with others in a spatial zone around themselves, which may be oriented either forwards or symmetrically around their centre, or with their nearest-neighbours or globally with all other individuals. Reported parameters are: , the baseline direction-change intensity; , the interaction radius for spatial models; , the number of interacting nearest-neighbours for topological models; and , the strength of interaction with negative and positive prawns respectively; and , the decay factor determining how long interaction effects persist.