Table 1.

Split -split- plot analysis of variance of tested environmental conditions for measured traits and group of species.

Variables	Species	Shade	Fertilization	Disturbance	S×F	S×D	F×D
RGRh	Invasive	0.012	0.001	0.471	0.016	0.964	0.334
	Early sc.	0.169	<0.001	0.447	0.312	0.610	0.204
	Late sc.	0.504	0.001	0.528	0.085	0.834	0.593
RSR	Invasive	0.161	<0.001	0.012	0.390	0.186	0.178
	Early sc.	0. 962	0.018	0.762	0.461	0.588	0.097
	Late sc.	0. 015	<0.001	0.066	0. 016	0.680	0.849
TLA	Invasive	0.065	0.007	0.017	0.084	0.559	0.215
	Early sc.	0.156	0.001	0.297	0.067	0.770	0.898
	Late sc.	0.017	0.001	<0.001	0.021	0.127	0.099
SLA	Invasive	< 0.001	0.168	0.115	0.249	0.023	<0.001
	Early sc.	< 0.001	0.020	0.253	0.976	0.720	0.776
	Late sc.	0.001	0.052	0.655	0.165	0.988	0.593
LWR	Invasive	0.012	<0.001	<0.001	0.021	0.383	0.965
	Early sc.	0.184	<0.001	0.077	0.268	0.965	0.801
Amax	Late sc.	0.437	<0.001	<0.001	0.034	0.784	0.349
	Invasive	0.710	0.043	0.168	0.450	0.897	0.986
	Early sc.	0.110	0.407	0.041	0.600	0.573	0.417
	Late sc.	0.588	0.005	0.008	0.055	0.243	0.553
Amaxw	Invasive	0.023	0.086	0.130	0.242	0.541	0.752
	Early sc.	0.002	0.800	0.512	0.771	0.986	0.947
	Late sc.	0.098	0.004	0.095	0.013	0.457	0.243
Nm	Invasive	0.836	<0.001	0.012	0.008	0.219	0.141
	Early sc.	0.603	<0.001	0.459	0.002	0.101	0.371
	Late sc.	0.037	<0.001	0.972	<0.001	0.773	0.548
PNUE	Invasive	0.253	0.213	0.171	0.398	0.048	0.107
	Early sc.	0.629	0.090	0.029	0.257	0.634	0.213
	Late sc.	0.056	0.037	0.023	0.120	0.777	0.426
Na	Invasive	<0.001	0.006	0.018	0.007	0.445	0.957
	Early sc.	<0.001	<0.001	0.479	<0.001	0.028	0.374
	Late sc.	<0.001	<0.001	0.872	<0.001	0.812	0.654

Significant p values (p < 0.05) are presented in bold. Species are grouped by strategy: the invasive species is Acer negundo, early-successional native species are Salix alba and Populus nigra and late-successional native species are Fraxinus excelsior and Fraxinus angustifolia. Traits are: RGR_h relative growth rate in seedling height, RSR root/shoot ratio, TLA total leaf area, SLA specific leaf area, LWR leaf weight ratio, A_max light-saturated assimilation rate per unit leaf area, A_maxw light-saturated assimilation rate per unit leaf dry weight, N_m nitrogen content, N_a nitrogen content per unit leaf area and PNUE the photosynthetic nitrogen use efficiency.