Table 1.

Empirical quantities required to test the transmission advantage and reproductive assurance hypotheses in nature

Term	Component	Parameters required to estimate component
δ = 1 – (Ws/Wx)	Inbreeding depression	Ws = lifetime fitness of selfed offspring
		Wx = lifetime fitness of outcross offspring
(px – ps)/y	Pollen discounting rate	ps = number of outcrossed seeds sired by selfing morphs
		px = number of outcrossed seeds sired by outcrossing morphs
(xx – xs)/y	Seed discounting rate	xs = number of outcrossed seeds made by selfing morphs
		xx = number of outcrossed seeds made by outcrossing morphs
		y = number of seeds produced by selfing

Both hypotheses require estimates of inbreeding depression and pollen discounting. The transmission advantage hypothesis assumes that seed discounting is complete, so this quantity is relevant only to the reproductive assurance hypothesis.