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TTL genes are required for male gametophytic development. Pollen and siliques from wild-type (WT) and ttl1/ttl2/ttl3/TTL4/ttl4 plants were analyzed by light microscopy. ttl1/ttl2/ttl3/TTL4/ttl4 plants showed abnormal, shrunken pollen grains (indicated by arrowheads). ttl1/ttl2/ttl3/TTL4/ttl4 plants do not support normal seed development, showing approximately 50% aborted seeds. [See online article for color version of this figure.]
