F |
number of farm fish in the area |
P |
total number of lice on farmed fish in the area |
|
average abundance of lice per farmed fish, P/F
|
L |
number of free-living copepodites from lice on farmed fish |
|
equilibrium values of , respectively |
L0
|
number of free-living copepodites from lice on wild fish |
μ |
louse mortality rate not caused by treatment of harvest (one per day) |
Tg
|
grow-out time (farmed fish harvest rate is ) (days) |
h = h1 + h2
|
louse mortality rate from harvest and treatment, respectively (one per day) |
|
scaled louse abundance used in §5. |
γ |
mortality rate of free-living copepodites (one per day) |
β |
transmission coefficient (captures per copepodite per host per unit time) |
λ |
natality (copepodites produced per adult female louse per unit time) |
ψ |
settlement success (probability an attached copepodite survives to adult) |
λ0, ψ0, μ0, γ0, β0, h0
|
reference values for λ, ψ, μ, γ, β, h, respectively |
|
scaled parameter (), and similarly for
|
|
the limit of at reference values () |
Fx
|
critical stocking level, given by equation (3.2) |
Fx0
|
Fx evaluated at λ0, ψ0, μ0, β0, γ0, h0
|
S |
salinity in parts per thousand (‰) |
θ |
temperature in °C |
aμ, aγ, aβ, aλ, aψ
|
salinity coefficients of μ, γ, β, λ, ψ (1/‰) |
bμ, bγ, bβ, bλ, bψ
|
temperature coefficients of μ, γ, β, λ, ψ (1/°C) |
gm, gstd |
geometric mean and standard deviation |
G |
weight of market fish (kg) |
Kx
|
critical production level (kT yr–1) |