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. 2011 Dec 22;287(11):7990–8000. doi: 10.1074/jbc.M111.315473

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FIGURE 1. — Genes of unknown function in SL-1 locus may be involved in biosynthesis. Genes associated with SL-1 biosynthesis are clustered in the M. tuberculosis genome, but one putative operon contains two genes of unknown function, sap (rv3821) and chp1 (rv3822). After sulfation of trehalose (Tre) by Stf0, T2S is successively esterified by PapA2 and PapA1. MmpL8 participates in the final two esterifications by an unknown mechanism and is essential for sulfolipid transport; Chp1 and Sap may also be involved.