Table 1.

Haplotype frequencies in a random sample

Locus A	Locus B		Allele frequency
	Allele 1	Allele 2
Allele 1	π11 = Qρ + QR(1 − ρ)	π12 = (1 − ρ)Q(1 − R)	Q = π11 + π12
Allele 2	π21 = (R − Q)ρ + R(1 − Q)(1 − ρ)	π22 = (1 − R)ρ + (1 − Q)(1 − R)(1 − ρ)	1 − Q = π21 + π22
Allele frequency	R = π11 + π21	1 − R = π12 + π22	1