Numerical solutions of evolutionary change in a weedy species growing
in a spatially implicit habitat in which fitness is limited both by
dispersal ability and by competitive ability, based on a model of
phenotypic diffusion (36). (A) Given this trade-off, an
initially weedy species, species 1, undergoes evolutionary change, with
its peak shown moving to the right. (B) After 50,000
years, species 1 has evolved into a much better competitor, but a much
poorer disperser than it originally was, and a new species, species 2,
has appeared. Species 2 is a superior disperser, but an inferior
competitor. It survives in vacant sites in this spatial habitat.
(C) Species 1 and 2 each evolve toward being superior
competitors. After some time a third species appears that is a poor
competitor, but excellent disperser. This third species evolves into a
superior competitor and a fourth species appears, etc. Shown here is
the result after 475,000 years, at which time 21 peaks of abundance
appear, each peak representing a different phenotype, thus
corresponding with different species.