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Known RNA-catalyzed reactions that are relevant to nucleotide biosynthesis. (A) Formation of 4-thiouridylate from free 4-thiouracil and ribozyme-tethered 5-phosphoribosyl-1-pyrophosphate. (B) 5′-phosphorylation of an oligonucleotide using γ-thio-ATP as the phosphate donor. (C) Activation of a nucleoside 5′-phosphate by formation of a 5′,5′-pyrophosphate linkage. (D) Template-directed ligation of RNA driven by release of a 5′,5′-pyrophosphate-linked adenylate.
