. 2012 May;191(1):261–277. doi: 10.1534/genetics.111.136440

Table 3. Expected offspring phenotype as a function of their own genotype and the genotype of their mother when there is a genomic imprinting effect.

	Offspring genotype
	A₁A₁	A₂A₁	A₁A₂	A₂A₂	Maternal litter average
Maternal genotype
A₁A₁	+a_m + a_o	+a_m + d_o – i_o	—	—	+a_m + a_op₁ + d_op₂ – i_op₂
	$(p_{1}^{3})$	*$(p_{1}^{2} p_{2})$*
A₁A₂, A₂A₁	d_m + a_o	d_m+ d_o – i_o	d_m+ d_o + i_o	d_m – a_o	+ d_m + $\frac{1}{2}$ d_o+ $\frac{1}{2}$ (a_o + i_o) (p₁ – p₂)
	$(p_{1}^{2} p_{2})$	$(p_{1} p_{2}^{2})$	$(p_{1}^{2} p_{2})$	$(p_{1} p_{2}^{2})$
A₂A₂	—	—	–a_m+ d_o + i_o	–a_m – a_o	– a_m – a_op₂ +d_op₁ + i_op₁
			*$(p_{1} p_{2}^{2})$*	$(p_{2}^{3})$
Offspring average	a_mp₁ + d_mp₂ + a_o	a_mp₁ + d_mp₂ +d_o – i_o	–a_mp₂ + d_mp₁+d_o+ i_o	–a_mp₂ + d_mp₁ – a_o

The format matches Table 1, except that, for the offspring, the reciprocal heterozygotes are separated into two classes that differ in the parent of origin of their alleles (with the paternally inherited allele listed first). The cells with dashes are the ones that would be used in the identification of additive maternal effects in the statistical cross-fostering design (Figure 1). The imprinting effect in these cells is boxed in boldface type.