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Expression of the smc02178-lacZ reporter gene fusion in M. sativa nodules. Nodules were observed at 7 dpi (A–E) and 14 dpi (F–J) after inoculation with S. meliloti 1021 (A and F), clr (B and G), cyaD1D2K (C and H), cyaK (D and I), and cyaD1D2 strains (E and J) carrying pGD2178.