Table 1. Analysis of pathogenic CNVs across multiple diagnostic categories: intellectual ability, Autism Spectrum Disorders, schizophrenia and bipolar disorder.
We analyzed 11 CNV loci where replicated evidence of association with schizophrenia and/or developmental disorders is available from multiple large-scale studies. We compared the frequency of each CNV loci in four major categories of neuropsychiatric disorders including intellectual disability (ID)/developmental delay (DD)/congenital malformations (CM), autism spectrum disorders (ASD), schizophrenia (SD) and bipolar disorder (BD)/Recurrent Depression, with its frequency in a large combined set of controls. We did not attempt a formal meta-analysis, as some studies did not include CNV data on both cases and controls. We present results with a pooled analysis of available CNV frequency data from different studies by performing a simple Fisher’s Exact Test and reporting odds ratio. We excluded overlapping individuals in different studies before doing tests for individual loci. Formal meta-analyses results are available for some of the published loci (Levinson et al., 2011) and the odds ratios (OR) and p-values reported for loci in the previous studies are similar to results reported here. Exonic deletions at the NRXN1 gene, also implicated as a risk factor for schizophrenia, were not analyzed as the CNVs affecting them are usually non-recurrent and much smaller than 500 kb and can be missed with the lower-resolution array studies included in the current analysis. See Supplemental Information for the references in the table.
CNV Locus | Position (Mb) | Disease category | Frequency in combined cases | Frequency in combined controls | P value | Odds Ratio [95% CI] | Case References | Control References |
---|---|---|---|---|---|---|---|---|
1q21.1 deletion | 145.0–146.3 | MR/DD/CM | 0.27% (102/ 38,330) | 0.02% (16/ 75,505) | 1.9 × 10−32 | 12.6 [7.4–21.3] | Mefford et al 2008; Brunettipierri et al 2008; Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2011; Grozeva et al 2010; Rucker et al 2011; Sanders et al 2011; Levinson et al 2011; Cooper et al 2011 |
ASD | 0.03% (1/ 3032) | 0.02% (16/ 75,505) | 0.49 | 1.6 [0.2–11.7] | Bucan et al 2009; Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.17% (21/ 12,174) | 0.02% (16/ 75,505) | 1.3 × 10−9 | 8.1 [4.3–15.6] | Vacic et al 2011; Levinson et al 2011 | |||
BD & Recurrent Depression | 0.02% (2/ 8264) | 0.02% (16/ 75,505) | 0.69 | 1.1 [0.3–5.0] | Malhotra et al 2011; Priebe et al 2011; Grozeva et al 2010; BiGS; Rucker et al 2011 | |||
1q21.1 duplication | 145.0–146.3 | MR/DD/CM | 0.14% (55/ 38,330) | 0.03% (19/ 57,730) | 2.7 × 10−9 | 4.4 [2.6–7.4] | Mefford et al 2008; Brunetti-pierri et al 2008; Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2010; Rucker et al 2011; Sanders et al 2011; Levinson et al 2011; Cooper et al 2011 |
ASD | 0.26% (8/ 3032) | 0.03% (19/ 57,730) | 3.6 × 10−5 | 8.0 [3.5–18.4] | Bucan et al 2009; Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.14% (13/ 9365) | 0.03% (19/ 57,730) | 0.0002 | 4.2 [2.1–8.6] | Vacic et al 2011; Levinson et al 2011 | |||
BD & Recurrent Depression | 0.10% (7/ 7382) | 0.03% (19/ 57,730) | 0.02 | 2.9 [1.2–6.9] | Malhotra et al 2011; Grozeva et al 2010; BiGS; Rucker et al 2011 | |||
3q29 deletion | 197.2–198.4 | MR/DD/CM | 0.07% (20/ 30465) | 0.002% (1/ 63,649) | 2.3 × 10−9 | 41.8 [5.6–311.6] | Ballif et al 2008; Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2011; Grozeva et al 2010; Sanders et al 2011; Levinson et al 2011; Mulle et al 2010 |
ASD | 0.05% (1/ 2120) | 0.002% (1/ 63,649) | 0.06 | 30.0 [1.9–480.4] | Sanders et al, 2011; Pinto et al 2010 | |||
SCZ | 0.10% (10/ 10,123) | 0.002% (1/ 63,649) | 2.3 × 10−8 | 63.0 [8.1–491.7] | Vacic et al 2011; Levinson et al 2011; Mulle et al 2010 | |||
BD | 0.04% (2/ 4659) | 0.002% (1/ 63,649) | 0.01 | 27.3 [2.5–301.5] | Malhotra et al 2011; Grozeva et al 2010; BiGS | |||
7q11.23 WBS deletion | 72.4–73.8 | MR/DD/CM | 0.27% (42/15767) | 0% (0/ 16,257) | 1.2 × 10−13 | Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2010; Sanders et al 2011; Cooper et al 2011; BiGS | |
ASD | 0% (0/2120) | 0% (0/ 16,257) | 1 | Sanders et al, 2011; Pinto et al 2010 | ||||
SCZ | 0% (0/8701) | 0% (0/ 16,257) | 1 | Vacic et al 2011; Levinson et al 2011; | ||||
BD | 0% (0/4659) | 0% (0/ 16,257) | 1 | Malhotra et al 2011; Grozeva et al 2010; BiGS | ||||
7q11.23 WBS duplication | 72.4–73.8 | MR/DD/CM | 0.10% (16/15767) | 0% (1/ 16,257) | 0.0001 | 16.5 [2.2–124.5] | Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2010; Sanders et al 2011; Cooper et al 2011; BiGS |
ASD | 0.19% (4/2120) | 0% (1/ 16,257) | 0.0008 | 30.7 [3.4–275.1] | Sanders et al, 2011; Pinto et al 2010 | |||
SCZ | 0.05% (4/8701) | 0% (1/ 16,257) | 0.053 | 7.5 [0.8–66.9] | Vacic et al 2011; Levinson et al 2011; | |||
BD | 0% (0/4659) | 0% (1/ 16,257) | 1 | Malhotra et al 2011; Grozeva et al 2010; BiGS | ||||
VIPR2 (7q36.3) duplication | 158.4–158.8 | MR/DD/CM | 0.06% (11/ 18,499) | Sanders et al 2011; Levinson et al 2011; Grozeva et al 2010. | ||||
ASD | 0.13% (3/ 2234) | 0.06% (11/ 18,499) | 0.18 | 2.3 [0.63–8.1] | Sanders et al 2011; Vacic et al 2011 | |||
SCZ | 0.19% (14/ 7336) | 0.06% (11/ 18,499) | 0.006 | 3.2 [1.5–7.1] | Levinson et al 2011 | |||
BD | 0.04% (2/ 4659) | 0.06% (11/ 18,499) | 1 | Malhotra et al 2011; Vacic et al 2011; Grozeva et al 2010 | ||||
15q11.2 deletion | 20.3–20.6 | MR/DD/CM | 0.51% (180/ 35,353) | 0.27% (201/ 75,486) | 4.9 × 10−10 | 1.9 [1.6–2.3] | Doornbos et al 2009; Mefford et al 2009; Burnside et al 2011; Cooper et al 2011 | Vacic et al 2011; Sanders et al 2011; Grozeva et al 2011; Rucker et al 2011; Cooper et al 2011; Kirov et al 2009; Levinson et al 2011 |
ASD | 0.09% (2/ 2120) | 0.27% (201/ 75,486) | 0.19 | 0.3 [0.1–1.4] | Pinto et al 2010; Sanders et al 2011 | |||
SCZ | 0.57% (72/ 12,665) | 0.27% (201/ 75,486) | 2.2 × 10−7 | 2.1 [1.6–2.8] | Vacic et al 2011; Kirov et al 2009; Levinson et al 2011 | |||
BD & Recurrent Depression | 0.22% (18/8264) | 0.27% (201/ 75,486) | 0.49 | Priebe et al 2011; Malhotra et al 2011; BiGS; Grozeva et al 2010; Rucker et al 2011 | ||||
15q11.2-13.1 duplication including PWS critical region | 20.8–26.2 | MR/DD/CM | 0.17% (27/ 15767) | 0.009% (5/ 53,832) | 2.2 × 10−13 | 18.5 [7.1–47.9] | Cooper et al 2011 | Vacic et al 2011; Cooper et al 2011; Glessner et al 2009; Sanders et al 2011; Ingason et al 2011 |
ASD | 0.39% (17/ 4315) | 0.009% (5/ 53,832) | 1.1 × 10−15 | 42.6 [15.7–115.5] | Glessner et al 2009; Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.05% (4/ 8384) | 0.009% (5/ 53,832) | 0.02 | 5.1 [1.4–19.1] | Vacic et al 2011; Ingason et al 2011 | |||
BD | 0% (0/ 4659) | 0.009% (5/ 53,832) | 1 | Malhotra et al 2011; Grozeva et al 2010; BiGS | ||||
15q13.3 deletion | 28.7–30.2 | MR/DD/CM | 0.26% (68/ 25,647) | 0.02% (13/ 74,106) | 6.1 × 10−28 | 15.1 [8.4–27.4] | Van bon et al 2009; Sharp et al 2008; Miller et al 2009; Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2011; Grozeva et al 2010; Rucker et al 2011; Sanders et al 2011; Cooper et al 2011; Levinson et al 2011 |
ASD | 0.19% (4/ 2120) | 0.02% (13/ 74,106) | 0.001 | 10.8 [3.5–33.1] | Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.19% (22/ 11,689) | 0.02% (13/ 74,106) | 2.1 × 10−11 | 10.7 [5.4–21.3] | Vacic et al 2011; Levinson et al 2011 | |||
BD & Recurrent Depression | 0.01% (1/ 7382) | 0.02% (13/ 74,106) | 1 | Malhotra et al 2011; Grozeva et al 2010; Rucker et al 2011 | ||||
16p13.11 duplication | 15.4–16.2 | MR/DD/CM | 0.30% (99/ 32,413) | 0.13% (81/ 62,973) | 8.4 × 10−9 | 2.4 [1.8–3.2] | Ramalingam et al 2011; Nagamani et al 2011; Mefford et al 2009; Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2011; Rucker et al 2011; Sanders et al 2011; Cooper et al 2011; Ingason et al 2009 |
ASD | 0.19% (4/ 2120) | 0.13% (81/ 62,973) | 0.36 | 1.5 [0.5–4.0] | Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.25% (13/ 5147) | 0.13% (81/ 62,973) | 0.03 | 2.0 [1.1–3.5] | Vacic et al 2011; Ingason et al 2009 | |||
BD & Recurrent Depression | 0.07% (6/ 7382) | 0.13% (81/ 62,973) | 0.38 | 0.8 [0.4–1.9] | Malhotra et al 2011; BiGS; Grozeva et al 2010; Rucker et al 2011 | |||
16p11.2 deletion | 29.5–30.2 | MR/DD/CM | 0.41% (64/ 15767) | 0.04% (25/ 56,752) | 7.3 × 10−24 | 9.2 [5.8–14.7] | Cooper et al 2011 | Grozeva et al 2011; Glessner et al 2009; Rucker et al Sanders et al 2011; Cooper et al 2011; Levinson et al 2011 |
ASD | 0.42% (18/ 4315) | 0.04% (25/ 56,752) | 2.0 × 10−10 | 9.5 [5.2–17.4] | Glessner et al 2009; Sanders et Pinto et al 2010 | |||
SCZ | 0.04% (4/ 9890) | 0.04% (25/ 56,752) | 1 | 0.9 [0.3–2.6] | Levinson et al 2011 | |||
BD & Recurrent Depression | 0.07% (6/ 8427) | 0.04% (25/ 56,752) | 0.28 | 1.6 [0.7–3.9] | Priebe et al 201; Mccarthy et al 2009; Rucker et al 2011 | |||
16p11.2 duplication | 29.5–30.2 | MR/DD/CM | 0.18% (18/ 15,767) | 0.03% (19/ 56,752) | 2.2 × 10−8 | 3.4 [1.8–6.5] | Cooper et al 2011 | Grozeva et al 2011; Glessner et al 2009; Rucker et al Sanders et al 2011; Cooper et al 2011; Levinson et al 2011 |
ASD | 0.39% (17/ 4315) | 0.03% (19/ 56,752) | 6.2 × 10−11 | 11.8 [6.1–22.7] | Glessner et al 2009; Sanders et Pinto et al 2010 | |||
SCZ | 0.31% (31/ 9890) | 0.03% (19/ 56,752) | 3.2 × 10−14 | 9.4 [5.3–16.6] | Levinson et al 2011 | |||
BD & Recurrent Depression | 0.13% (11/ 8427) | 0.03% (19/ 56,752) | 0.0008 | 3.9 [1.9–8.2] | Priebe et al 2011; Mccarthy et al 2009; Rucker et al 2011 | |||
17p12/HNPP deletion | 14–15.4 | MR/DD/CM | 0.02% (3/ 15767) | 0.02% (14/ 59,086) | 1 | 0.8 [0.2–2.8] | Cooper et al 2011 | Vacic et al 2011; Cooper et al 2011; Grozeva et al 2011; Sanders et al 2011; Kirov et al 2009 |
ASD | 0.09% (2/ 2120) | 0.02% (14/ 59,086) | 0.1 | 4.0 [0.9–17.5] | Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.14% (8/ 5891) | 0.02% (14/ 59,086) | 0.0004 | 5.7 [2.4–13.7] | Vacic et al 2011; Kirov et al 2009 | |||
BD | 0.05% (3/ 5541) | 0.02% (14/ 59,086) | 0.17 | 2.3 [0.7–8.0] | Priebe et al 2011; Malhotra et al 2011; BiGS; Grozeva et al 2010 | |||
17q12 deletion | 31.9–33.2 | MR/DD/CM | 0.10% (32/ 31,516) | 0.006% (4/ 68,131) | 1.4 × 10−12 | 17.3 [6.1–49.0] | Cooper et al 2011; Moreno-deluca 2010 | Vacic et al 2011; Grozeva et al 2011; Sanders et al 2011; Moreno-deluca 2010; Cooper et al 2011. |
ASD | 0.09% (2/ 2120) | 0.006% (4/ 68,131) | 0.01 | 16.0 [2.9–87.9] | Sanders et al 2011; Pinto et al 2011 | |||
SCZ | 0.06% (4/ 7142) | 0.006% (4/ 68,131) | 0.004 | 9.5 [2.4–38.2] | Vacic et al 2011; Moreno-deluca 2010 | |||
BD & Recurrent Depression | 0% (0/ 4659) | 0.006% (4/ 68,131) | 1 | Malhotra et al 2011; BiGS; Grozeva et al 2010 | ||||
22q11.21 deletion | 17.1–18.7 | MR/DD/CM | 0.61% (96/ 15767) | 0% (0/ 70,739) | 8.4 × 10−72 | Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2011; Rucker et al 2011; Sanders et al 2011; Cooper et al 2011; Levinson et al 2011 | |
ASD | 0.07% (2/ 3032) | 0% (0/ 70,739) | 0.002 | Bucan et al 2009; Sanders et al 2011; Pinto et al 2010 | ||||
SCZ | 0.30% (36/ 12,202) | 0% (0/ 70,739) | 1.0 × 10−30 | Vacic et al 2011; Levinson et al 2011 | ||||
BD & Recurrent Depression | 0.05% (4/ 7382) | 0% (0/ 70,739) | 8.0 × 10−5 | Malhotra et al 2011; Grozeva et al 2010; BiGS; Rucker et al 2011 | ||||
22q11.2 duplication | 17.1–18.7 | MR/DD/CM | 0.32% (50/ 15767) | 0.08% (23/ 27,133) | 5.9 × 10−8 | 3.7 [2.3–6.1] | Cooper et al 2011 | Vacic et al 2011; Grozeva et al 2010; Sanders et al 2011; Cooper et al 2011; Glessner et al 2009; ISC et al 2008; Rucker et al 2011 |
ASD | 0.28% (12/ 4315) | 0.08% (23/ 27,133) | 0.002 | 3.3 [1.6–6.6] | Glessner et al 2009; Sanders et al 2011; Pinto et al 2010 | |||
SCZ | 0.03% (3/ 8701) | 0.08% (23/ 27,133) | 0.17 | 0.4 [0.1–1.4] | Vacic et al 2011; MGS; ISC et al 2008 | |||
BD & Recurrent Depression | 0.05% (4/ 7382) | 0.08% (23/ 27,133) | 0.49 | 0.6 [0.2–1.8] | Malhotra et al 2011; Grozeva et al 2010; BiGS; Rucker et al 2011 |