Table 2.

Model variables

Variable	Symbol	Unit	Equation
Individual length	L(i, t)	cm	[3, 5, 7, 9]
Individual weight	W(i, t)	kg	[6]
Individual fecundity	Q(i, t)	–	[6]
Offspring number	n0(i, t)	–	[11]
Genotypic growth coefficient	g(i)		[2, 8]
Genotypic PMRN intercept	y(i)	cm
Genotypic PMRN slope	s(i)	cm·year−1	[1]
Genotypic gonado-somatic index	r(i)	–
Phenotypic growth coefficient	G(i, t)		[2, 5]
Phenotypic PMRN intercept	Y(i)	cm
Phenotypic PMRN slope	S(i)	cm·year−1	[1]
Phenotypic gonado-somatic index	R(i)	–	[5, 6]
Instantaneous predation mortality rate	mp	year−1	[7, 8]
Instantaneous foraging mortality rate	mf	year−1	[8]
Maximum instantaneous harvest rate	fmax	year−1	[10]
Instantaneous harvest mortality rate	F(i, t)	year−1	[10]
Total instantaneous mortality rate	Z(i, t)	year−1

–, dimensionless variables.